Moult and Ageing of European Passerines

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A brand-new, completely revised second edition of Jenni and Winkler's classic guide, updated and improved for the next generation of ringers and professional ornithologists.

Lukas Jenni and Raffael Winkler have studied moult across a wide range of bird species for decades, and in this book bring their observations together to produce a valuable reference for both professional ornithologists and bird ringers. This second edition has been completely updated and revised, with 16 new species accounts added, bringing the total covered to 74.

The first part of the book provides an up-to-date summary of the moult strategies and moult sequences of European passerines, and discusses the ecological consequences of moult. Throughout the book, the authors draw on the enormous amount of data on moult that they have collected over 40 years of study and which, combined with data from the literature, allow them to present a thorough synthesis of the subject.

The second part is of particular value to ringers. Following a general introduction to ageing, detailed moult profiles are given for 74 European passerine species, illustrating all of the major moult strategies and including useful summary statistics, schematic diagrams of the extent of moult and indications of the variation within each species. The main moult strategies are illustrated with schematic graphs, and the moult strategies and extent of moult of every European passerine species are summarised in tabular form.

The crowning feature of this book is its collection of more than 600 full-colour photographs of extended wings, which show the entire range of moult patterns and plumage-ageing criteria. An appendix gives supplementary information on ageing birds by the degree of pneumatisation of the skull.

Large in format, packed with high-quality photography and lavish in production specifications, this second edition of
Moult and Ageing in European Passerines is both a major reference for ornithologists, zoologists, bird ringers and dedicated birdwatchers, and a work of great scholarship and beauty.

Author(s): Lukas Jenni, Raffael Winkler
Edition: 2
Publisher: Helm
Year: 2020

Language: English
Pages: 336
City: London

Cover
Half-title
Title
Copyright
Contents
Preface
Part I
Chapter 1. The function and consequences of moult
1.1 Functions of the plumage
1.2 Plumage maintenance and the need for plumage renewal
1.2.1 Feather maintenance and wear
1.2.2 Adjustments to the plumage
1.3 The processes and costs of moult
1.3.1 The products and processes of moult
1.3.2 Costs of moult
1.3.3 Conclusions
1.4 Environmental effects on new feather quality
1.5 Fitting moult into the annual cycle
Chapter 2. Terminology and methods in moult research
2.1 Arrangement of the feathers
2.1.1 Flight-feathers
2.1.2 Wing-coverts
2.2 Terminology of plumages, feather generations and moults
2.2.1 Concepts of moult and plumage terminologies
2.2.2 General terms
2.2.3 Moult terms
2.2.4 Terms for plumages, feathers and feather generations
2.2.5 Age classes
2.3 Recording moult, plumage and feather properties
2.3.1 What to record: moult progress or moult intensity and of which feather tract?
2.3.2 Scoring flight-feather moult
2.3.3 Scoring body-feather moult
2.3.4 Mass of feathers
2.3.5 Quantification of feather wear
2.3.6 Feather growth rates
2.3.7 Physical properties of feathers and fault bars
2.4 Analysing moult data
2.4.1 Seasonal timing and duration (progress) of moult
2.4.2 Moult intensity and moult speed
2.4.3 Sequence of moult
2.4.4 Extent of moult
Chapter 3. The moult of adults
3.1 Introduction to the moult strategies
3.2 Sequence of moult
3.2.1 Basic sequence of the complete moult
Flight-feathers
Body-feathers and wing-coverts
3.2.2 Functional aspects of the basic sequence of moult
3.2.3 Variations and exceptions to the basic moult sequence
Primaries
Secondaries
Tertials
Rectrices
Wing-coverts
Variation in the relationships between flight-feather tracts
3.3 Moult strategies
3.3.1 Complete post-breeding moult in the breeding area: Moult strategies 1 and 2
Complete post-breeding moult in the breeding area
Partial pre-breeding moult
3.3.2 Complete moult in the non-breeding area: Moult strategy 3
Partial moult before autumn migration
Suspension of the complete moult within the non-breeding area
Additional partial pre-breeding moult
Conclusions
3.3.3 Seasonally divided moult of remiges: Moult strategy 4
Seasonally divided primary moult (moult strategy 4a)
Seasonally divided secondary moult (moult strategies 4b and 4c)
3.3.4 Partial and complete biannual moult of remiges: Moult strategy 5
3.3.5 Summary and concluding remarks
3.4 Timing and duration of the complete moult
3.4.1 Timing and duration of the complete post-breeding moult in the breeding area
Timing of moult
Moult duration
3.4.2 Moult under time constraints
Reduction of moult duration
Overlap between breeding and moult
Overlap between moult and autumnal activities
Arrested moult
Transfer of moult to the non-breeding season
3.4.3 Timing of moult in trans-Saharan migrants
3.5 Summary and concluding remarks
3.6 Summary graphs and table of the moult strategies of European passerines
Chapter 4. The moult during the first year of life
4.1 Introduction
4.2 The juvenile plumage
4.2.1 Development of the juvenile plumage and completion after fledging
4.2.2 Structure and quality of the juvenile plumage
4.2.3 Coloration of the juvenile plumage
4.3 Sequence of post-juvenile moult
4.3.1 Sequence of the complete post-juvenile moult
4.3.2 Sequence of the partial post-juvenile moult
General sequence
Wing-coverts and alula
Tertials
Secondaries
Rectrices
4.3.3 Eccentric and other sequences of partial primary moults (post-juvenile and first pre-breeding)
4.3.4 Sequence and priorities of moult
4.4 How juveniles assume the adult moult strategies
4.4.1 Moult strategies 1 and 2: partial (complete) post-juvenile and complete post-breeding moult in the breeding area
Partial pre-breeding moult
4.4.2 Moult strategies 3 and 5: complete moult in the non-breeding area
Incomplete first pre-breeding moult
4.4.3 Moult strategy 4: seasonally divided moult
First partial pre-breeding moult including primaries
First partial pre-breeding moult including secondaries
4.4.4 Summary and concluding remarks
4.5 Variation in extent, timing and duration of the post-juvenile moult in the breeding area
4.5.1 Experimental evidence of control of post-juvenile moult
4.5.2 Intraspecific variation in timing and extent of the partial post-juvenile moult
Variation with hatching date
Differences between populations
Differences between the sexes and effects of energetic and nutrient stress
Intraspecific variation in the extent of post-juvenile moult during the course of the non-breeding season and between wintering sites
4.5.3 Interspecific variation in timing and extent of the partial post-juvenile moult
Timing and duration
Extent
4.5.4 Partial post-juvenile primary moult in the breeding area
4.5.5 Complete post-juvenile moult in the breeding area
4.6 Summary and concluding remarks
Part II
Chapter 5. Ageing European passerines
5.1 Ageing criteria in live birds
5.2 Ageing using plumage characters
5.2.1 Recognition of juvenile feathers
Structure and shape
Coloration
Wear
Growth bars and fault bars
5.2.2 Differences in extent of moult
5.2.3 Differences between post-juvenile and subsequent feather generations
5.3 General ageing criteria in European passerines based on moult
5.3.1 Species with a complete post-juvenile moult in the first summer/autumn: Moult Type A
5.3.2 Species with a partial post-juvenile/complete post-breeding moult in the breeding area: Moult Type B
5.3.3 Species with a partial post-juvenile/complete post-breeding moult in the breeding area and a partial pre-breeding moult in winter/spring: Moult Type C
5.3.4 Species with a complete moult in the non-breeding area: Moult Type D
Chapter 6. Species accounts
6.1 Presentation of the data and how to use it
6.1.1 Material
6.1.2 Relevance of the data
6.1.3 Presentation and analysis of the data
6.2 Procedure of ageing
Species accounts
Lanius collurio Red-backed Shrike
Lanius senator Woodchat Shrike
Lanius nubicus Masked Shrike
Oriolus oriolus Eurasian Golden Oriole
Garrulus glandarius Eurasian Jay
Pica pica Eurasian Magpie
Nucifraga caryocatactes Spotted Nutcracker
Corvus corone Carrion Crow
Bombycilla garrulus Bohemian Waxwing
Periparus ater Coal Tit
Cyanistes caeruleus Eurasian Blue Tit
Parus major Great Tit
Riparia riparia Sand Martin
Hirundo rustica Barn Swallow
Delichon urbicum Common House Martin
Phylloscopus trochilus Willow Warbler
Phylloscopus collybita Common Chiffchaff
Acrocephalus schoenobaenus Sedge Warbler
Acrocephalus scirpaceus Eurasian Reed Warbler
Acrocephalus palustris Marsh Warbler
Hippolais icterina Icterine Warbler
Locustella naevia Common Grasshopper Warbler
Locustella fluviatilis River Warbler
Sylvia atricapilla Eurasian Blackcap
Sylvia borin Garden Warbler
Sylvia nisoria Barred Warbler
Sylvia curruca Lesser Whitethroat
Sylvia communis Common Whitethroat
Troglodytes troglodytes Eurasian Wren
Sitta europaea Eurasian Nuthatch
Sturnus vulgaris Common Starling
Turdus torquatus Ring Ouzel
Turdus merula Common Blackbird
Turdus pilaris Fieldfare
Turdus iliacus Redwing
Turdus philomelos Song Thrush
Turdus viscivorus Mistle Thrush
Muscicapa striata Spotted Flycatcher
Erithacus rubecula European Robin
Luscinia svecica Bluethroat
Luscinia luscinia Thrush Nightingale
Luscinia megarhynchos Common Nightingale
Ficedula hypoleuca European Pied Flycatcher
Phoenicurus ochruros Black Redstart
Phoenicurus phoenicurus Common Redstart
Saxicola rubetra Whinchat
Oenanthe oenanthe Northern Wheatear
Cinclus cinclus White-throated Dipper
Passer domesticus House Sparrow
Passer montanus Eurasian Tree Sparrow
Prunella modularis Dunnock
Motacilla flava Western Yellow Wagtail
Motacilla cinerea Grey Wagtail
Motacilla alba alba White Wagtail
Anthus campestris Tawny Pipit
Anthus pratensis Meadow Pipit
Anthus trivialis Tree Pipit
Anthus spinoletta Water Pipit
Fringilla coelebs Common Chaffinch
Fringilla montifringilla Brambling
Coccothraustes coccothraustes Hawfinch
Pyrrhula pyrrhula Eurasian Bullfinch
Chloris chloris European Greenfinch
Linaria cannabina Common Linnet
Acanthis cabaret Lesser Redpoll
Loxia curvirostra Red Crossbill
Carduelis carduelis European Goldfinch
Carduelis citrinella Citril Finch
Serinus serinus European Serin
Spinus spinus Eurasian Siskin
Emberiza citrinella Yellowhammer
Emberiza cia Rock Bunting
Emberiza hortulana Ortolan Bunting
Emberiza schoeniclus Common Reed Bunting
Appendix: The use of skull pneumatization for ageing
The process of skull pneumatization
Recognition of skull pneumatization
Skull pneumatization scores
Age determination using skull pneumatization
Explanations of the graphs
References
Scientific names with their English, German, French, Italian and Spanish translations
Index
Quick reference key